Collaboration Meetings

Ethogram for Amphiprion percula

Adapted from Wong et al. 2013 & Pacaro et al. 2023

Behavioural Category	Behaviour	Description
Social affiliation	Meeting	Male and female simultaneously approach each other to within 1 body length (may be followed by soft touch)
Social affiliation	Soft touch	Male and female make contact with fins or body before separating again, preceded by meeting
Aggressive	Chase	Fish rapidly swims toward a conspecific and the conspecific flees
	Bite	Fish bites another fish
	Aggressive display	Fish orient themselves toward an aggressor
	Ram	Fish rams head-first into an opponent
Submissive	Flee	Fish rapidly swims away after being chased or bitten
	Quiver	Fish shakes body side-to-side in a jolty motion (may serve submissive function) (Body shake in Wong 2013)
	Submissive display	Fish meets another, turns side-on with erect fins and stiffened posture
	Body shake	Fish shakes body side-to-side toward female (context-dependent signal)
Maintenance	Feed	Fish orients toward food and consumes particles
	Carry	Fish digs and spits out sand or debris
	Bite anemone	Fish bites the mouth or surface of the anemone
	Yawn	Fish opens mouth slowly and widely
	Bathing	Fish swims slowly and deliberately through tentacles
	Clearing	Fish removes debris from the anemone base using vigorous fin movements
	Massaging	Fish interacts with or attends to the anemone
Reproductive	Mouthing	Fish uses mouth to pick at eggs
	Fanning	Energetic pectoral fin movement beside egg clutch
	Tending	Time individual spends with eggs
	Nest preparation	Fish cleans algae near the base of the anemone
	Vigorous mouthing	Forceful mouthing where parents appear to bite and fling their eggs away from the clutch with their mouth
	Vigorous fanning	Energetic form of fanning where parents use their entire body to fan, rather than their pectoral fins
Other	Dart	Sudden quick swim in any direction, not directed at anything (Jolt in Wong 2013)
Other	Undefined	Behaviors that are not defined or unclear